meaning - "Changes in" , "Changes of" or "Changes to"


I am confused about the selection of in, of or to I want to explain that "changes in hydrological variables and changes in landscape variables in wetlands can change the populations of waterbirds" Here are some options, I am not sure they are right or not. Can someone please help that which one is explaining it best and why?


1) Changes in wetland hydrological and landscape variables can alter the population dynamics of waterbirds.


2) Changes of hydrology and landscapes in wetlands can alter the population dynamics of waterbirds.


3) Changes to hydrology and landscapes in wetlands can alter the population dynamics of waterbirds.


4) Hydrological and landscape changes in wetlands can alter the population dynamics of waterbirds.




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